A plant growing in the wrong place is sometimes described as a weed, but it can be more than just an inconvenience.  Plants in the wrong place can have a devastating effect on ecosystems, dramatically reducing biodiversity.  The same is true for certain animals - those that have 'traveled' from their natural habitats to 'foreign areas’ where they have no natural predators to keep their numbers in check.  Foreign or alien species are sometimes introduced to an area in the belief that they will solve local problems with pests or over-population of a native species.   A classic example of this is the cane toad.  Native to  parts of central and South America, the cane toad is preyed upon by a variety of animals such as caimans, snakes, eels and some fish species.  It was introduced into sugar plantations in various parts of the world to control beetles that were damaging sugar cane crops. Since this was a successful strategy in Puerto Rico, the toad was introduced elsewhere, notably Australia.  Just over a hundred toads were released in Queensland in 1935, with more released in 1937.  Unfortunately the toad did not effectively control the grey-backed cane beetles (the intended target), and they found other things to eat.  Their numbers grew exponentially and they spread into other areas, such as the Northern Territory and New South Wales.  These poisonous toads have significantly reduced biodiversity particularly affecting native amphibians and reptiles. Sometimes foreign species are introduced as ornamental plants or 'exotics'. During the Victorian period many plant species were brought to the UK for country estates.  Rhododendrons that were collected across Asia and the Himalayas became popular in parks and gardens.  However, Rhododendron ponticum, a species native to the southwestern Iberian Peninsula (parts of Portugal and Spain) and the southern Black Sea Basin (parts of Bulgaria & Turkey)  has been less well received.   It was introduced in the eighteenth century by Conrad Loddiges.  Loddiges grew seed and sold on young plants as an ornamental flowering shrubs for gardens, parks and estates. On country estates and heathland areas, it was planted as cover for game birds as its dense growth offered shelter and protection. It has since spread aggressively and is now considered to be an invasive species. An invasive species is a non-native species that spreads and damages its new environment. R. ponticum poses a threat to key woodland ecosystems, such as Atlantic Oak Woodland.  When this shrub ‘invades’, it comes to dominate the area.  It creates deep shade so the woodland floor becomes a dark and ‘barren’ place. Many ground flora species are lost so that only shade tolerant mosses and liverworts remain.  These plants form a ‘dense mat’ of vegetation that is a barrier to seed germination.  Additionally, there is evidence that as R. ponticum grows, it produces chemicals which inhibit the growth of other species.  This phenomenon is known as allelopathy.  Even when the Rhododendron is removed, it is difficult to reestablish the original flora.  Clearing an area of this plant is difficult and expensive. One effective method involves drilling the stems and injecting herbicide directly into the plant, a targeted approach that uses a lot less herbicide than spraying.  Mabberley’s Plant-book notes that the cost of eradicating the plant from Snowdonia was £30M and that was in 1988. Another Victorian introduction was the grey squirrel. Native to North America, grey squirrels were first released in the UK in the latter part of the 19th century.   One person associated with their spread was the 11th Duke of Bedford  (Herbrand Russell).   He was involved in various animal conservation projects, but he released and 'gifted' many grey squirrels from the estate at Woburn.  The populations of grey squirrels in Regent's Park, London are thought to have come from there. Humans may still be helping the spread of the grey squirrel, albeit unintentionally.  One squirrel was captured on the Isle of Skye (in 2010), it had traveled from Glasgow, as revealed by its genetic profile.  It had probably made the journey as stowaway under a car bonnet. It is important that we are aware of how 'easy' it is for these animals to travel with us.  Their introduction has been disastrous for the native red squirrel populations, due to competition and the spread of the squirrel pox virus.  It is vital that grey squirrels do not colonise areas where the red squirrel still survives. . More recent introductions have arrived due to increased global trade.  Parasites and pests can ‘hitch a ride’ with people, materials or goods as they move across the world.   Climate change is also altering the range and distribution of many plant and animal species.  There is an alert for the Asian or Yellow Legged hornet (Vespa velutina nigrithorax), an invasive, non-native species from Asia.  This hornet preys upon honeybees and other beneficial insects such as hover flies and bumblebees. It hovers outside bee hives, waiting to catch and then kill returning honeybees. The effect on bee colonies can be devastating. The Asian Hornet arrived in France in 2004 (through an import of Chinese pottery) and has spread rapidly. Now it is found in Italy, Spain, Portugal, Switzerland, Belgium and Germany.  Preventing its establishment in the UK is critical.  Any sighting needs to be reported, this can be done through the Asian Hornet Watch. This link downloads a PDF, which gives more information about the Asian Hornet, and contains images comparing the  appearance of the asian hornet, the european hornet, the wasp and honey bee. Other invasive species that are a cause for concern include : Himalayan Balsam Harlequin ladybirds, Signal Crayfish, Eight-toothed bark beetle Citrus long horned beetle Giant Hogweed Thanks to Anton for images.

Though some insects are problematic in that they are carriers of disease (for example, mosquitos and malaria, ticks and Lyme disease); it is nevertheless true that without insects food chains and ecological systems would collapse.  Insects act as  pollinators not just of garden flowers, but of crops,  natural pest control agents (ladybirds eat aphids), Decomposers, breaking down waste products of other animals, remains of dead plants and animals (dung beetles). Their activity ensures the recycling of nutrients in complex biogeochemical cycles. However, as the woodlands’ blog has reported previously, insects (like so many wildlife species) are under threat.  They are in decline due to loss and damage of habitats,  climate change,  pollution and  pesticide use. The decline in insect numbers (in the UK) is ‘monitored’ through BUGLIFE and Kent Wildlife Trust.  Each year a survey is undertaken using the ‘splatometer technique’, in which motorists are asked to record the number of flying insects (e.g. moths, flies, aphids, bees and flying beetles) that are squashed on their front number plate (after a journey).  The length of the journey is recorded, a photo taken and count details uploaded via the BUGLIFE APP, the app includes a tutorial and some safety advice.  Comparing this year’s results of over 6000 journeys with those gathered in 2004 (by the RSPB, who used the same method) reveals a dramatic fall in flying insect numbers.   London, for example, showed a dramatic fall in numbers of 91%.  The fall across England was 83%, Wales saw a 79% drop and Scotland a 76% fall.   Whilst figures for Northern Ireland were limited they suggest a 54% decline. [caption id="attachment_21589" align="alignleft" width="300"] bumblebees favour teasels[/caption] Insects [and other wildlife species] can be helped by: Creating larger areas of natural habitats (many have been lost to roads, agriculture, urban expansion) Creating wildlife corridors to join up similiar habitats/ecosystems throughout the landscape Creating wild flower ‘meadows’ by road sides, verges etc Reducing the use of pesticides and other chemicals which have significant effects on wildlife.  The effect of neonicotinoids on bees and bumblebees is well documented.       

Many natural ecosystems are periodically exposed to fire.  After a fire, there is often reduced competition and increased nutrient availability (from ash etc.).  The plants and flowers that grow after a fire are visited more often by pollinators, such as bees and other insects.  This can result in increased production of fruits and seeds. Bushfires have been part of certain australian ecosystems for thousands of years and some native species are ‘fire adapted’.  They have come to 'rely' on fires as a means of reproduction and / or  dispersal. Whilst no one fire can be attributed to climate change alone, rising temperatures and aridity, lengthening of the ‘fire season’, combined with bursts of extreme ‘fire weather’, all combine to suggest climate change is implicated. As the frequency of fires increases, the possible benefits of fire to such ecosystems / species are being lost. Fire can help with the physical dispersal of seeds from the parent plant.  In some parts of the world, such as South Africa and Australia, fire and / or smoke can be the stimulus for seed dispersal and subsequent germination.  Plants such as some species of Protea, Banksia, certain members of the myrtle family (e.g. some Eucalypts), and some Pines and Sequoias 'make use' of fire to disperse their seeds. Seed dispersal involving fire is termed serotiny.  Many of these plants produce woody fruits or cones in which the seeds are held.  The mechanism underlying seed release varies but can be due to a resin that ‘seals’ the seeds inside the fruit or cone.  The resin ‘melts’ / liquefies on exposure to heat releasing the seed or there may be a structure called a seed separator (as in Banksia).  Serotinous conifers (like lodgepole pine), have mature cones in which the cone scales are naturally sealed shut with resin.   Most of the seeds stay in the canopy until the cones reach 122-140o F  (i.e 50 to 60oC).  At these temperatures, heat / fire  melts the resin and  the cone scales open to expose the seed. The seed can then drop or drift to a burned but cooling ash-rich soil bed. The seeds do well on the burnt soil available to them as the site offers reduced competition, more light, warmth plus the nutrients from the burning of leaves and litter.  Some species align their germination to immediate post-fire conditions - stimulated by chemicals present in the smoke.  The organic compounds karrikins,  products of the degradation of cellulose are  a germination ‘cue’ for some species.  Karrikins are thought to be present on the soil surface after a fire.  When it rains,  the karrikins are 'washed' into the soil, and seeds present in the soil seed bank are then stimulated to germinate. Thanks to Steve Sangster and John Cameron for images of woodland fire.  

The end of lockdown, and the peace that came with it, was what made me want some natural solitude as the world got busy again. Having a share of a near 100-acre ancient wood should provide that given there are no public footpaths through it, a locked gate and farm land all around. It was late summer when I ‘got the keys’ and I was recovering from major surgery, so over the autumn into winter I’ve pottered and observed.  Each visit the place looks different as the leaves fall, different fungi come and go, and the wood is deluged by each storm. Storm Babet blew down the largest silver birch tree. The kids discovered it – wading through the thick brown bracken in a clearing I’d yet to explore. They had fun running along it. So far, no more have come down, and I’m glad I bought the Beech Tree wood owners insurance before the first storms hit. There is about one acre of older pine that stand majestic, almost as a guard for the younger trees beyond. We’ve re-planted our Christmas tree here. I hope it takes! Then it’s a grassy break, that allows machinery to manage a drainage dyke, across a wooden bridge and into the 4 acres of dense young birch coppice. This needs a lot of thinning out, if owt else is to grow there. The west boundary is a huge sheep field and the sun pours in. It wasn’t until my fifth visit that I managed to get through to a far corner of the plot and find another small clearing, where clearly deer had been laying on the dead bracken. I hesitate to say basking in the winter sun ….....  in Yorkshire! There’s evidence of badger setts too but none seem active. Woodcock abound and I will have to control the dog in spring. It isn’t silent but I’m surprised there isn’t a greater dawn chorus when I visit. Perhaps the buzzards and red kites are scaring smaller birds off.   I certainly hear those birds of prey about. Is this a sign of climate change or a lack of tree diversity? To help the both I’ve planted some cuttings of hazel and walnut, and a variety of seeds – conkers, acorns, sloes, sweet chestnuts, sycamore and walnut, but goodness knows if anything will germinate, or whether the squirrels have had a feast!  It was a rushed affair as I wasn’t completely recovered from surgery when I did it. There are lots of plots within the greater wood and I’ve met many of my neighbours, who are all very friendly. At least one is an outstanding wildlife expert and I’m sure there’s lots to learn from them.  I don’t have great plans as yet other than increasing the diversity of trees. A fig tree at home next to a west facing wall does really well.   Will the west facing aspect enable similar here?  There seems to be some quite large structures within the wider wood. On my plot, there’s a small hut and I don’t have plans for anything more. I just want to observe for a year, open up a couple of paths through the jungle and create somewhere flat to camp. There is wonderful solitude and the dog loves it – even though she has come back with ticks. Neighbours have told me their 'tick stories' and I’ve noted that they wear thick overalls and boots.   Probably best  if I string up a hammock to watch the sunsets!  

Tree survival and drought. Researchers at the University of California have been working on a method that helps predict whether forests / woodlands can survive periods of drought.  As climate change is altering patterns of snow and rainfall, so periods of drought are likely to become more common. Forests are important in terms of carbon sequestration, that is, they take up carbon dioxide from the air and convert it into sugars, starches etc that are stored in the leaves, branches, stems and roots.  However, in order to assimilate and convert carbon dioxide (in photosynthesis), trees (indeed all plants) need a supply of water.  When water is limited, trees need to make use of their reserve materials.  Just as we make use of body reserves of fat and glycogen when food / diet in inadequate. However, reserves can only sustain a tree for a finite period of time.  If drought persists, the tree reaches a ’tipping point’ and it will die.  The researchers studied a forest in the Sierra Nevada that experienced a period of drought between 2012 and 2015.   During this period, millions of trees died.  The team recorded rainfall, soil moisture and temperature in the forest AND the amount of carbon dioxide that the trees absorbed, and their reserve materials.  They found that the trees were able to maintain function / health after the onset of the drought but with the passing of time, the trees exhausted their reserves and were unable to use / convert carbon dioxide into food.  They had reached the tipping point and died.   The methodology of this study was called CARDAMON (carbon data assimilation with a model of carbon assimilation); it is hoped that it can be used to evolve strategies to enhance forest and woodland resilience in the face of climate change. Pollinators. [caption id="attachment_35902" align="aligncenter" width="675"] hoverfly[/caption] University researchers from the UK and Finland have been trying to determine the most effective pollinators of crop plants, like strawberries (and other fruits).  Plentiful and effective pollinators are needed to ensure a good harvest of the fruits. The researchers studied the pollinators at three strawberry farms through the (long) growing season for the fruit.  They adopted two approaches : They caught the insects that visited the strawberry flowers and analysed the pollen they carried in detail (pollen load and type). They also counted the number of flower visits by the different insects, (a quick way to identify key local pollinators). Many insects were identified, including :-  European drone fly :           Eristalis arbustorum Honeybee :                               Apis mellifera Levels drone fly :                   Eristalis abusivus Buff tailed bumblebee :     Bombus terrestris White tailed bumblebee :  Bombus lucorum Common drone fly :             Eristalis tenax Red tailed bumblebee :      Bombus lapidarius Early bumblebee :                Bombus pratorum Bent-shinned Morellia :   Morellia aenescens Hoverflies are true flies, that is, they belong to the order Diptera or true flies, as they have a pair of wings and a pair of halteres (balancing  / orienteering organs used when in flight). Several of the flies in the genus Eristalsis are known as Drone Flies (due to their resemblance to honey bee drones).  The larvae of Eristalis  species are commonly found in putrid / stagnant water and sometimes referred to as “rat-tailed maggots”. It was noted that pollinators also made use of the wild plants to supplement their diets, as strawberries alone cannot meet the nutritional needs of pollinators.  ‘Elsanta’ strawberries have a relatively low sucrose and protein content in both their nectar and pollen. The precise  order of importance of pollinators varied between farms.  Bee (Apis and Bombus) species  and hoverfly (Eristalis) emerged as key pollinators. The European drone fly was the most important pollinator at two of the three farms studied, evidence that hoverflies can be effective pollinators.  One farm had commercial hives of the honey bee but they were less significant than the activities of of the hoverflies and bumblebees. The abundance of a particular insect, coupled with its active period were /  are important determinants of pollinator importance.  Sawdust and plastics - a possible use?. Plastics represent a relatively new, but persistent and major form of pollution (on land, in the sea, indeed everywhere).  Whilst many plastic objects are instantly visible in the form of discarded bottles, fast food containers, many plastic pollutants are in the form of very small particles of plastics  - nano and microplastics.  The concern is that we and other organisms are taking these microscopic particles into our bodies from our food / drinking water. However, it is possible that plant materials may offer some ‘solutions’.  Water that contains micro and nano plastics can be filtered through sawdust that has been treated with tannic acid.   Tannic acid is large molecule, its molecular formula is C72H52O46 .  Tannic Acid is found in certain plant galls (swelling of trees caused by parasitic wasps) and in the twigs of certain trees, such as Chestnut and Oak.  The wood sawdust contains fibres of cellulose, combined with hemicelluloses and lignin.  Water can flow through this material by capillary action.  This plant-based filtration (known as bioCap) of plastic-laden water is capable of dealing with a wide range of nanoplastics (PVC, PET, polyethylene etc), and tests with mice suggest that the filtered water may be sufficiently free of plastic to pose little risk.  

Scotland’s west coast has a number of temperate rain woodlands / forests. They are quite rare. The remnants of oak, birch, ash, native pine and hazel woodlands are small and isolated from each other. They are noteworthy for the diversity and richness of the bryophytes (mosses and liverworts) and lichens; found in abundance on the trees, rocks and on the ground.   Sadly, such woodlands have been in decline for some time. In the past, this woodland covered large areas of the west coast of Scotland, but much has been lost over the last two thousand years.  These woodlands / forests now cover a small area, just under 5% of the land. Factors that have contributed to the decline and loss of this woodland include:- mismanagement,  overgrazing by sheep and  invasion by non-native species [such as Rhododendron ponticum]. According to recent study by Scottish Environment LINK, deer now represent a considerable threat to the woodlands.  Whilst deer are part of woodland ecosystems, when their numbers increase beyond a certain point then they become a significant problem.  Deer numbers are now at historic highs in Scotland/  Money has been made available to manage surging deer populations, for example, through the provision of deer fencing.  However, the report considers that such fencing is “both expensive and often ultimately ineffective”.  More needs to be done if deer damage is to be reduced and allow regeneration of the woodlands. Developing a community approach to deer stalking and management will be important, combined with the use of technologies such as thermal and drone surveying. A greater focus on the management of roe and sika deer, combined with the removal of Rhododendron ponticum will be needed if the woodlands are to flourish and expand. see also : https://www.thescottishfarmer.co.uk/news/23637346.soaring-deer-numbers-see-new-powers-land-managers/   [caption id="attachment_39688" align="aligncenter" width="675"] Rhododendron ponticum, these plants were growing near the River Tay.[/caption] visit https://www.instagram.com/woodlands.co.uk/?hl=en  

Some years back, the Woodlands blog posted various articles about hedgerows,  noting the loss of many - due to the increased mechanisation of farming in the mid C20th.  Now, there is greater recognition of the importance of hedgerows, and there are initiatives to promote the maintenance and expansion of hedgerows. But what is a hedgerow? Natural England offers a definition as follows : A hedgerow is defined as any boundary line of trees or shrubs over 20m long and less than 5m wide, and where any gaps between the trees or shrub species are less that 20m wide (Bickmore, 2002). Any bank, wall, ditch or tree within 2m of the centre of the hedgerow is considered to be part of the hedgerow habitat, as is the herbaceous vegetation within 2m of the centre of the hedgerow.  This differs from the definition in the  Biodiversity Action Plan, which included references to ancient hedges / species-rich hedges.  The definition now includes all hedgerows consisting of at least one native woody species of tree or shrub (mainly), but it does exclude bramble and honeysuckle as ‘woody species’.  According to one source, there are some 550,000km of hedgerow in England, with over 400,000 km being actively managed.  Hedgerows are an important semi-natural habitat in what is otherwise a managed agricultural landscape. They are found across the country but there are more in lowland regions. Hedgerows in the south east are associated with large fields and fewer trees, the proportion of trees in hedgerows increases as one goes north and west.   The nature of hedgerows varies across the country but all are important as : They provide a habitat, shelter (micro-climate provision) and resources for many different species (from plants to insects, birds and mammals). Hedgerows are particularly important as nesting sites for birds. They support animals that have key roles within the broader ecosystem, for example pollinators and predators of pests. They offer an important source of nectar that helps support wild bees - adjacent farmland can be a poor source of nectar Hedgerows are known to support threatened (red listed) species Hedgerows capture and store carbon (above and below ground) Hedgerows offer ecosystem services eg. mitigation of water flux and availability, landscape connectivity, soil conservation / stabilisation. A number of studies indicate that increasing the number of hedgerows would help with landscape connectivity (for example, for hedgehogs) and that planting of blackthorn and hawthorn in association with later flowering species would help support a number of wild bee species.  Expanding the number of hedgerows could have some negative effects as they might offer a home to invasive species and / or pathogens; but one study has indicated that ash trees in hedgerows suffer less impact from ash dieback than trees in forests.  To date there does not appear to be any detailed research on whether increasing hedgerow coverage would have any impact on tree disease / pathogen spread. Hedgerows, like woodlands themselves, face a number of challenges due to climate change.  Warmer winters may mean that the ‘winter chill’ requirements of some shrubs / trees will not be met; this may mean flowers and fruits fail to form properly which in turn means less food for birds, small mammals etc.  Drier summers may stress some species, trees like Beech are susceptible to drought.  Extreme weather events (like Storm Arwen) can inflict damage on hedgerow trees.  If a hedgerow is next to farmland, then it may experience drift from pesticide and / or herbicide spraying  nutrient enrichment (eutrophication) due to the use of fertilisers. Hedgerows with a diverse structure, with plants, shrubs and trees of varying ages and heights provide the widest range of niches / microhabitats for wildlife.  The inclusion of dead / decaying wood offers opportunities for various fungi, saproxylic beetles, woodlice etc.  Some hedgerows are managed / reduced with a mechanical flail (see above !!!). If this is done annually, it can result in a loss of biodiversity. Trimming should be done on a 2 or 3 year cycle; and some sections of the hedge might be left for longer " see (https://www.hedgelink.org.uk/cms/cms_content/files/76_ne_hedgecutting.pdf).  Thousands of tree and hedgerow plants are being planted to create a flood defence project at Castlehill, East Hull.   The plan is to create some seven hectares of woodland and over five kilometres of new hedgerow, as part of a flood defence project (to store floodwater east of the city).  Trees such as field maple, downy birch, English oak, and black alder are being planted along with species of willow, dog rose, guelder rose and blackthorn and hawthorn to create hedgerows and scrubland.  Other species will be allowed to naturally develop in the area and the habitat is expected to reach ‘maturity in some fifteen to twenty years. There is a citizen science project that involves surveying hedgerows.  It is organised by the People’s Trust for Endangered Species [PTES].  The Great British Hedgerow Survey guidelines can be found here : https://hedgerowsurvey.ptes.org/survey-guidelines Some times hedges offer a home to other things         

In the eighteenth and nineteenth century, many explorers / adventurers brought ‘exotic’ plants back to the United Kingdom. These ‘exotics’ were planted in arboreta, botanic gardens, and some in gardens.  One bush that was introduced was Rhododendron ponticum. The plant is native to the eastern and western Mediterranean, and the Pontic Mountains, hence its name). It was introduced into Britain in the late eighteenth century, by Conrad Loddiges.  It was planted in Victorian hunting estates, also on heathland areas to provide shelter for game species. Its rootstock has been used for grafting of less hardy, more colourful types.  Many Rhododendron species are a delight and an adornment to our parks and gardens, indeed many hundreds of species of Rhododendron are known (many in China and the Himalayas)*.   Rhododendron ponticum has proved to be invasive.  It is a threat to key parts of our woodland ecosystems, such as Atlantic Oak Woodland.  Atlantic oak woodland is sometimes referred to as Celtic Rainforest.  It is characterised by lichen covered trees, growing amongst a rich moss and liverwort flora.  This woodland environment is damp and humid, to which streams and waterfalls contribute. These woodlands have evolved under the influence of the Gulf Stream,  which helps keeps warm and wet the area.  In some parts of the country, the woodlands have remained in their 'ancient state', since the last ice age.  However, these woodlands were more extensive but now exist as much smaller ‘pockets’ - damaged by grazing, pollution, and ‘exotics or aliens’ like R. ponticum. When this shrub ‘invades’, it 'takes over' and the woodland floor becomes a dark and barren place. A deep shade results from the thickets of the Rhododendron.  This results in the loss of much of the ground flora so that only some shade tolerant mosses and liverworts remain.  They form a ‘mat’ of dense vegetation that is a barrier to seed germination.  Even when the Rhododendron thicket is removed, the re-establishment of the original flora is compromised.  There is also evidence that as it grows this shrub produces chemicals which inhibit the growth of other species;  this is known as allelopathy. R.ponticum has spread in many areas, mainly to the West of the country. Each flower can produce several thousand seeds, so that a large bush can produce several million seeds in year. These seeds are tiny and wind dispersed; and though not all the seeds will germinate and grow, many will and colonise an area. Even when bushes have been removed from an area and the litter layer cleared, the seeds may persist in the seed bank of the soil - allowing the species to recolonise. In consequence, follow up over a five year period is really important. Recent figures suggests that some 37,000 hectares are affected in the UK.  Though the government does make a grant available for the removal of Rhododendron, progress with its removal has been slow. In Wales, there is a project called the Celtic Rainforests Project  (YouTube video link here) that focuses on invasive species and their attempted eradication in Atlantic Oak Woodlands in Wales.  With the agreement of the landowner, the project will organise surveys to identify the scale of the problem, and then contractors to carry out the work over the period of a management agreement, at no cost to the landowner [woodlands.co.uk has groups of buyers who have agreed for their various woods to be covered by such management agreements]. Clearing an area of this plant is difficult and expensive. An  effective first treatment to eliminate R. ponticum is to drill the stems, and inject herbicide directly into the plant.  This uses a lot less herbicide than spraying, and is a selective approach.  Mabberley’s Plant-book notes that the cost of eradicating the plant from Snowdonia was £30 M and that was in 1988.  Further information about the work in the welsh oak woodlands here. The plant is also a problem in Ireland. Indeed, referring to the Killarney National Park a politician has said “nothing short of calling in the army is going to put it right.” [caption id="attachment_39688" align="aligncenter" width="675"] Rhododendron ponticum growing near to the River Tay.[/caption] Forest Research has a number of publications about the management and control of R. ponticum. * Details of the genus may be found in Mabberley's Plant-Book. With thanks to Chris Colley